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Circus is monophyletic and Accipiter is polyphyletic. Abstract The monophyly of the raptorial Circus genus harriers has never been in question, but the specific status of many, often vulnerable island endemic, taxa remains uncertain. Here we utilise one mitochondrial and three nuclear loci from all currently recognised Circus taxa species and subspecies to infer a robust phylogeny, to estimate the divergence date and to reconstruct the biogeographic origins of the Circus group. Our phylogeny supports both the monophyly of Circus and polyphyly of the genus Accipiter. Depending on the rate of molecular clock used, the emergence of the harrier clade took place between 4. The sister relationship of the Northern Harrier C. Thus both should be elevated to species status: Further, the African Marsh C. The remaining marsh harriers exhibit very little genetic diversity, and are all recently diverged taxa that exhibit allopatric distributions.

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Towns in the region. Another debris flow from Corona spread towards Salar de Atacama. Its large crater is filled by another cone, [53] which forms the highest summit of the Lascar volcano. The eastern cone also known as Activo [53] is capped off with three distinct craters [52] which are delimited by arcuate fractures. There are many fumaroles along the rim of the inner crater.

There are traces of a collapse of the volcano towards the northeast, with an associated horseshoe-shaped scar. The flow advanced just north of the head of Quebrada Talabre before passing over cliffs and entering it. Its rocks have a pale gray-blue colour, and their composition resembles the Soncor flow, despite more mafic lavas and pyroclastics being erupted in the time period between the emplacement of the Soncor flow and the Capricorn Lava. It was emplaced after the collapse of the oldest edifice, covering Aguas Calientes’ western slopes.

The flow deposit was later modified by glacial activity. On the western slope, it buries the even older Piedras Grandes flow, which crops out only at the margins of the Soncor flow. It was accompanied by a Plinian fall deposit. Finally, the andesitic pumice Tumbres flow is found on the northwest—west—southwestern slopes of Lascar.

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The Lyonnais dialect was partly replaced by the French language as the importance of the city grew. Lyon also features a renowned opera house. Many are to be seen everywhere around the city. This old tradition is now expending in a contemporary expression. See for example Guillaume Bottazzi art work. The African Museum of Lyon is one of the oldest museums situated in Lyon. In their designation, UNESCO cited the “exceptional testimony to the continuity of urban settlement over more than two millennia on a site of great commercial and strategic significance.

Gastronomy For several centuries Lyon has been known as the French capital of gastronomy , due, in part, to the presence of many of France’s finest chefs in the city and its surroundings e. This reputation also comes from the fact that two of France’s best known wine-growing regions are located near Lyon: Beaujolais wine is very popular in Lyon and remains the most common table wine served with local dishes. Lyon is the home of very typical and traditional restaurants: Bouchons are usually convivial restaurants serving local dishes, and local wines.

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